Publications – Ward Strong

This researcher no longer works for the B.C. Government.


Listed in order of publication date:

2016 to 2020:


  • Lesieur, V, E. Lombaert, T. Guillemaud, B. Courtial, W. Strong, A. Roques, M.-A. Auger-Rozenberg. 2019. The rapid spread of Leptoglossus occidentalis in Europe: a bridgehead invasion. Journal of Pest Science 92(1): 189-200.


  • Woods, Jack and Ward Strong. 2018. Impact of Surround®, Pounce®, and Matador® on Lodgepole Pine Filled Seed Production in Southern Interior BC Seed Orchards: 2017 Trial. Canadian Forest Genetics Association, Tree Seed Working Group Newsbulletin 67: 7-15.
  • Lesieur, V., E. Lombaert, T. Guillemaud, B. Courtial, W. Strong, A. Roques, M. A. Auger-Rozenberg. 2018. The rapid spread of Leptoglossus occidentalis in Europe: a bridgehead invasion. DOI: 10.1007/s10340-018-0993-x


  • Strong, Ward. 2016. New pesticide registrations for the Canadian seed orchard industry. Canadian Forest Genetics Association, Tree Seed Working Group Newsbulletin 63: 22-26.
  • Acheampong, S., Ward B. Strong, M.D. Schwartz, R.J. Higgins,  M.A. Thurston, E.J. Walker, and J. Roberts. 2016. First Canadian records for two invasive seed-feeding bugs, Arocatus melanocephalus (Fabricius, 1798) and Raglius alboacuminatus (Goeze, 1778), and a range extension for a third species, Rhyparochromus vulgaris (Schilling, 1829) (Hemiptera: Heteroptera). J Ent Soc BC 113:74-78.
  • Justin G. A. Whitehill, Hannah Henderson, Ward Strong, Barry Jaquish, and Jörg Bohlmann. 2016. Function of Sitka spruce stone cells as a physical defence against white pine weevil. Plant, Cell & Environment 39: 2545-2556.
  • Tamara A. Richardson , Ward B. Strong, Brian H. Aukema, Stephen Takàcs, Tracy Zahradnik and and B. Staffan Lindgren. 2016. Assessment of cues potentially mediating host selection of Leptoglossus occidentalis on Pinus contorta. Agriculture and Forest Entomology 19(3) 245-256.

2011 to 2015:


  • Strong, Ward. 2015. Thermocouples, iButtons, and Lepto-cams: understanding the low Pli seedset problem. Forest Genetics Council of British Columbia, TICTalk 12: 13-18.
  • Strong, Ward B. 2015. Lodgepole pine seedset increase by mesh bagging is due to exclusion of Leptoglossus occidentalis (Hemiptera: Coreidae). Journal of the Entomological Society of British Columbia 112: 3-17
  • Woods, Jack and Ward Strong. 2015. Impact of Matador on lodgepole pine filled seed production in Southern Interior BC seed orchards. Canadian Forest Genetics Association, Tree Seed Working Group Newsbulletin  61: 16-20.


  • Roversi, P. F., G. Sabbatini Peverieri, M. Maltese, P. Furlan, W. B. Strong and V. Caleca. 2014. Pre-release risk assessment of the egg-parasitoid Gryon pennsylvanicum for classical biological control of Leptoglossus occidentalis. Journal of Applied Entomology 138: 27–35


  • Strong, Ward. 2013. Lepto-cams and thermocouples: Why do insect bags prevent August Pli seed loss in the Interior of BC? Canadian Forest Genetics Association, Tree Seed Working Group Newsbulletin 57: 9-10.
  • Sabbatini Peverieri G., P. Furlan, D. Benassai, S. Caradonna, W.B. Strong, and P.F. Roversi. 2013. Host Egg Age of Leptoglossus occidentalis (Heteroptera, Coreidae) and Parasitism by Gryon pennsylvanicum (Hymenoptera, Platygastridae). Journal of Economic Entomology 106(2): 633-640.


  • Zaradnik, T., S. Takacs, W. Strong, R. Bennett, A. Kuzmin, and G. Gries. 2012. Douglas-fir cone gall midges respond to shape and infrared wavelength attributes of host tree branches. Can. Entomol. 144(5):658-666
  • Glenn P. Svensson, Hong-Lei Wang, Jean-Marc Lassance, Olle Anderbrant, Guo-Fa Chen, Berit Gregorsson, Claud Guertin, Eevamaria Harala, Erling V. Jirle, Ilme Liblikas, Vladimir Petko, Alain Roques, Olle Rosenberg, Ward Strong, Kaljo Voolma, Tiina Ylioja, Yan-Jun Want, Xiao-Ming Zhou, and Christer Löfstedt. 2012. Assessment of genetic and pheromonal diversity of the Cydia strobilella species complex (Lepidoptera: Tortricidae). Systematic Entomology 38(2): 305-315.
  • \Maltese,  Matteo, Virgilio Caleca, Emilio Guerrieri, and Ward B. Strong. 2012. Parasitoids of Leptoglossus occidentalis Heidemann (Heteroptera: Coreidae) recovered in Western North America and first record of its egg parasitoid Gryon pennsylvanicum (Ashmead) (Hymenoptera: Platygastridae) in California. Pan-Pacific Entomologist 88(3):347–355.
  • Peverieri, G.S., P. Furlan, S. Simoni, W.B. Strong, and P.F. Roversi. 2012. Laboratory evaluation of Gryon pennsylvanicum (Ashmead) (Hymenoptera, Platygastridae) as a biological control agent of Leptoglossus occidentalis Heidemann (Heteroptera, Coreidae). Biol. Control 61(1):104-111 


  • Roversi, P.F., W.B. Strong, V. Caleca, M. Maltese, G.S. Peverieri, L. Marianelli, L. Marziali, and A. Strangi. 2011. Introduction into Italy of Gryon pennsylvanicum (Ashmead), an egg parasitoid of the alien invasive bug Leptoglossus occidentalis Heidemann. EPPO Bulletin 41:72-75 
  • Whitehouse, C.M., A.D. Roe, W.B. Strong, M.L. Evenden, and F.A.H. Sperling. 2011. Biology and management of North American cone-feeding Dioryctria species. Can. Entomol. 143(1):1-34.

2006 to 2010:


  • Strong, W.B. and R.G. Bennett. 2010. Sampling plan for Adelges cooleyi (Hemiptera: Adelgidae) in spruce seed orchards. Can. Entomol. 142(1):14-23. 


  • Bains, B., F. Isik, W.B. Strong, B. Jaquish, J.A. McLean, and Y.A. El-Kassaby. 2009. Genetic resistance of spruce to gall-forming adelgids (Hemiptera: Adelgidae). Can. J. For. Res. 39(12):2536-2541. 


  • Strong, W.B., J.G. Millar, G.G. Grant, J.A. Moreira, J.M. Chong, and C. Rudolph. 2008. Optimization of pheromone lure and trap design for monitoring the fir coneworm, Dioryctria abietivorella. Entomologia Experimentalis et Applicata 126(1):67-77 
  • Gries, W.B. Strong, and R.G. Bennett. 2008. Vibratory communication signal produced by male western conifer seed bugs (Hemiptera: Coreidae). Can. Entomol. 140(2):174-183.
  • Takacs, S., H. Bottomley, I. Andreller, T. Zaradnik, J. Schwarz, R. Bennett, W. Strong, and G. Gries. 2008. Infrared radiation from hot cones on cool conifers attracts seed-feeding insects. Proc. R. Soc. B 276:649-655 


  • Bains, B., W. Strong, and J. McLean. 2007. Adelgid Research at Kalamalka Forestry Centre. TICtalk 8(1):29-32.
  • Strong, W. 2007. Cone and Seed Pest Research Report. TICtalk 8(1):24-28. 


  • Corrigan, J., M. Carlson, G. Giampa, V. Berger, C. Walsh, and W. Strong. 2006. They’re Heeeere! - Observations of the First Mountain Pine Beetle Attacks Recorded at the Kalamalka Forestry Centre. TICtalk 7(1):4-8.
  • Strong, W.B. 2006. Seasonal changes in seed reduction in lodgepole pine cones caused by feeding of Leptoglossus occidentalis (Hemiptera: Coreidae). Can. Entomol. 138(6):888-896. [Link]

2000 to 2005:


  • Millar, J.G., G.G. Grant, J.S. McElfresh, W. Strong, C. Rudolph, J.D. Stein, and J.A. Moreira. 2005. (3Z,6Z,9Z,12Z,15Z)-Pentacosapentaene, a Key Pheromone Component of the Fir Coneworm Moth, Dioryctria abietivorella. J. Chem. Ecol. 31(5):1229-1234. [Link]


  • Heeley, T., R.I. Alfaro, L. Humble, and W. Strong. 2003. Distribution and life cycle of Rhyacionia buoliana (Lepidoptera: Tortricidae) in the interior of British Columbia. J. Entomol. Soc. B.C. 100:19-25 
  • Sopow, S.L., J.D. Shorthouse, W. Strong, and D.T. Quiring. 2003. Evidence for long-distance, chemical gall induction by an insect. Ecology Letters 6(2):102-105 


  • Bates, S.L., W.B. Strong, and J.H. Borden. 2002. Abortion and Seed Set in Lodgepole and Western White Pine Conelets Following Feeding by Leptoglossus occidentalis (Heteroptera: Coreidae). Environ. Entomol. 31(6):1023-1029 


  • Strong, W.B., S.L. Bates, and M.U. Stoehr. 2001. Feeding by Leptoglossus occidentalis (Hemiptera:Coreidae) reduces seed set in lodgepole pine (Pinaceae). Can. Entomol. 133(5):857-865. 

1999 and earlier:


  • Carlson, M., W. Strong, C. Walsh, and A. Yanchuk. 1999. Breeding for white pine blister rust resistance: the Kalamalka white pine seed orchard. TICtalk 2(1):12-13.